Anisodactylus (Pseudoanisodactylus) signatus (Panzer, 1796)
Family: Carabidae
Size: 12.7 mm (10.0 to 13.5 mm)
Distribution: Transpalaeartcic, Spain to Siberia, Asia Minor, Japan, Korea
Biology: Eurytop
Location: Russia, Volgograd, Kamyshensky, Golubaja river valley, 150 m
leg. E.Alexeer, 15.-19.IV.2000; det. Wrase, 2016
Photo: U.Schmidt, 2017

Anisodactylus signatus   (Flickr)

Location List Coleoptera Russia  (Google Drive)

Location List Carabidae Palaearctic Region  (Google Drive)